1. Department of Biological, Geological, and Environmental Sciences (BiGeA), Laboratory of Molecular Anthropology and Centre for Genome Biology, University of Bologna, Bologna, Italy
2. School of Anthropology and Museum Ethnography, University of Oxford, Oxford, UK
3. Interdepartmental Center "L. Galvani," (CIG), University of Bologna, Bologna, Italy
4. IRCCS, Institute of Neurological Sciences of Bologna, Bologna, Italy
5. Department of Experimental, Diagnostic and Specialty Medicine (DIMES), University of Bologna, Bologna, Italy
6. Applied Biomedical Research Center (CRBA), S. Orsola-Malpighi Polyclinic, Bologna, Italy
7. Biodemography of Aging Research Unit, Social Science Research Institute, Duke University, Durham, NC 27708, USA
8. Istituto Auxologico Italiano IRCCS, Cusano Milanino, Milan, Italy
9. Department of Biology, Ecology and Earth Sciences, University of Calabria, Rende, Italy
10. Geriatric Unit, Department of Medical Sciences and Community Health, Milan, Italy
11. Fondazione Ca' Granda, IRCCS Ospedale Maggiore Policlinico, Milan, Italy
12. Department of Experimental and Clinical Biomedical Sciences "Mario Serio", University of Florence, Florence, Italy
13. Department of Neuroscience, Psychology, Drug Research and Child Health, University of Florence, Florence, Italy
14. IRCCS Don Gnocchi, Florence, Italy
15. Azienda Ospedaliera-IRCCS, Reggio Emilia, Italy
16. Department of Surgical, Medical, Dental and Morphological Sciences with Interest Transplant, Oncological and Regenerative Medicine, , Italy
17. Department for the Cultural Heritage (DBC), University of Bologna, Ravenna, Italy
18. Co-senior authors
19. Clinical Chemistry, Department of Laboratory Medicine, Karolinska Institutet at Huddinge University Hospital, S-141 86 Stockholm, Sweden
20. CNR Institute of Molecular Genetics, Unit of Bologna, Bologna, Italy
21. Rizzoli Orthopaedic Institute, Laboratory of Cell Biology, Bologna, Italy