1. Department of Cell Biology and Physiology, University of North Carolina, Chapel Hill, NC 27599, USA
2. Division of Gastroenterology, University of Pennsylvania, Philadelphia, PA 19104, USA
3. Department of Molecular Biomedical Sciences, College of Veterinary Medicine, North Carolina State University, Raleigh, NC 27695, USA
4. Department of Hematology, Stanford University, Stanford, CA 94305, USA
5. Massachusetts General Hospital and Harvard Medical School, Boston, MA 02115, USA