1. Key Laboratory of Bio-resources and Eco-environment, Ministry of Education, College of Life Science, Sichuan University, Chengdu 610064, China
2. Institute for Advanced Study, Chengdu University, Chengdu 610106, China
3. College of Life Sciences and Food Engineering, Yibin University, Yibin 644000, China
4. The Sichuan Key Laboratory for Conservation Biology of Endangered Wildlife, Chengdu Research Base of Giant Panda Breeding, Chengdu 610081, China