1. Department of Medicine, Division of Geriatrics, University of Utah, Salt Lake City, UT 84112, USA
2. Geriatric Research, Education, and Clinical Center, George E. Whalen VA Medical Center, Salt Lake City, UT 84112, USA
3. Department of Exercise and Sport Science, University of Utah, Salt Lake City, UT 84112, USA
4. CRMBM, Aix-Marseille Universite, CNRS 7339, Marseille, France
5. Department of Radiology and Utah Center for Advanced Imaging Research, University of Utah, Salt Lake City, UT 84112, USA