1. Department of OphthalmologyCase Western Reserve UniversityThe Research Institute of University Hospitals of Cleveland Cleveland Ohio 44106 USA
2. Department of NeurologyCase Western Reserve UniversityThe Research Institute of University Hospitals of Cleveland Cleveland Ohio 44106 USA
3. Department of NeurosciencesCase Western Reserve UniversityThe Research Institute of University Hospitals of Cleveland Cleveland Ohio 44106 USA
4. Department of The Comprehensive Cancer CenterCase Western Reserve UniversityThe Research Institute of University Hospitals of Cleveland Cleveland Ohio 44106 USA
5. Department of ToxicologyUniversity of Kentucky Lexington Kentucky 40536 USA
6. Department of Physiology and BiophysicsUniversity of Washington Seattle Washington 98195 USA
7. Max-Planck-Institute for Biochemistry Martinsried Germany
8. School of Biological SciencesWellcome Trust Centre for Cell-Matrix Research Manchester United Kingdom