1. Laboratory of Animal Genetics and Reproduction Research Faculty of Agriculture Hokkaido University Sapporo Japan
2. Laboratory of Theriogenology Faculty of Veterinary Medicine Hokkaido University Sapporo Japan
3. Laboratory of Animal Reproduction Department of Animal Science School of Veterinary Medicine Kitasato University Towada Japan
4. Global Station for Food, Land and Water Resources Global Institution for Collaborative Research and Education (GI‐CoRE) Hokkaido Japan
5. Livestock Research Institute Aomori Prefectural Industrial Technology Research Center Aomori Japan
6. NODAI Genome Research Center Tokyo University of Agriculture Setagaya Japan
7. Department of Embryology Nara Medical University Kashihara Japan
8. Department of Bioscience Tokyo University of Agriculture Setagaya Japan