1. Guangdong Provincial Key Laboratory of Marine Biotechnology, Shantou University, Daxue Road, 515063, Shantou, Guangdong, China
2. Institute of Tropical Aquaculture, Universiti Malaysia Terengganu, Mengabang Telipot, 21030, Kuala Terengganu, Terengganu, Malaysia
3. Division of Biological Sciences, College of Arts and Sciences, University of the Philippines Visayas, 5023, Miagao, Iloilo, Philippines