1. Graduate School of Fisheries and Environmental Sciences, Nagasaki University, Bunkyo-machi 1-14, Nagasaki, Nagasaki 852-8521, Japan;
2. Faculty of Fisheries, Nagasaki University, Bunkyo-machi 1-14, Nagasaki, Nagasaki 852-8521, Japan;
3. Sasebo Municipal Fisheries Division, Hachiman-cho 1-10, Sasebo, Nagasaki 857-8585, Japan