1. Department of Integrative Biology, Oregon State University, 3062 NW Snowberry Place, Corvallis, OR 97330
2. Department of Fisheries and Oceans Canada, Pacific Biological Station, 3190 Hammond Bay Road, Nanaimo, BC V9T 6N7, Canada
3. Department of Fisheries and Oceans, Institute of Ocean Sciences, 9860 West Saanich Road, Sidney, BC V8L 4B2, Canada