1. Key Laboratory of East China Sea and Marine Fishery Resource Exploitation and Utilization, Ministry of Agriculture, East China Sea Fisheries Research Institute, Chinese Academy of Fishery Science, 300 Jungong Road, Yangpu District, 200090 Shanghai, China
2. College of Marine Sciences, Shanghai Ocean University, 999 Huchenghuan Road, Pudong New District, 201306, Shanghai, China
3. Department of Fisheries, Animal and Veterinary Sciences, University of Rhode Island, 9 East Alumni Avenue, Kingston, RI 02881