1. Department of Marine Life Science (BK21 FOUR) and Marine Science Institute, Jeju National University, 102 Jejudaehak-ro, Jeju 63243, Republic of Korea
2. Tidal Flat Research Center, West Sea Fisheries Research Institute, National Institute of Fisheries Science (NIFS), 177-1 Sandannambuk-ro, Gunsan 54001, Republic of Korea
3. Jeju Fisheries Research Institute, National Institute of Fisheries Science, 6 Yeondaemaeul-gil, Jeju 63068, Republic of Korea