1. School of Marine Biosciences, Kitasato University, Minami-ku, Sagamihara, Kanagawa 252-0373, Japan
2. Faculty and Graduate School of Fisheries Sciences, Hokkaido University, Minato-cho, Hakodate, Hokkai
3. Southeast Asian Fisheries Development Center, Aquaculture Department (SEAFDEC/AQD), 5021 Tigbauan, I