1. Department of Forest Environment Protection, Kangwon National University, Chuncheon, 200-701, Korea
2. Department of Microbiology and Plant Pathology, Forestry and Agricultural Biotechnology Institute (FABI), University of Pretoria, Pretoria, 0002, South Africa
3. Division of Forest Insect Pests and Diseases, Korea Forest Research Institute, Seoul, 130-712, Korea