1. Microorganism Resources Division, National Institute of Biological Resources, Incheon 22689, Korea
2. School of Life Science, Kyungpook National University, Daegu 41566, Korea
3. COREANA Innovation Research Center, Cheonan 27002, Korea
4. Research Institute for Dok-do and Ulleung-do Island, Kyungpook National University, Daegu 41566, Korea