1. Laboratory of Reproductive Endocrinology, Department of Animal Reproduction, National Institute of Animal Industry, Ministry of Agriculture, Forestry and Fisheries, Ibaraki 305-0901, Japan
2. Tokyo Research Laboratories, Kyowa Hakko Kogyo Co., Ltd., Tokyo 194-8533, Japan
3. Department of Degenerative Neurological Diseases, National Institute of Neuroscience, National Center of Neurology and Psychiatry, Tokyo 187-8502, Japan