1. Department of Veterinary Pathology, Infection and Public Health, Institute of Veterinary Science, University of Liverpool, Neston, United Kingdom.
2. Istituto Zooprofilattico Sperimentale delle Venezie, Legnaro, Italy.
3. Chester Zoo, Chester, United Kingdom.
4. Department of Infection Biology, Institute of Infection and Global Health, University of Liverpool, Neston, United Kingdom.