1. Center for Tsukuba Advanced Research Alliance, University of Tsukuba, Tennoudai, Tsukuba, Ibaraki, Japan
2. GeneCare (previously AGENE) Research Institute, Kamakura, Kanagawa Japan
3. Department of Rheumatology, Tokyo Metropolitan Otsuka Hospital, Minami-Otsuka, Toshima-ku, Tokyo, Japan
4. Tennoudai, Tsukuba, Ibaraki, Japan
5. Department of Cellular and Molecular Biology, Hiroshima University School of Medicine, Minami-ku, Hiroshima, Japan