1. CAS and Shandong Province Key Laboratory of Experimental Marine Biology, Center for Ocean Mega-Science, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China.
2. Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou), Guangzhou, China.
3. Laboratory for Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology, Qingdao, China.
4. University of Chinese Academy of Sciences, Beijing, China.
5. Qingdao Agricultural University, Qingdao, China.