1. Department of Otolaryngology–Head and Neck Surgery, Graduate School of medicine, Kyoto University, Kyoto City, Japan.
2. Department of Virus Research, Institute for Life and Medical Sciences, Kyoto University, Kyoto City, Japan.
3. Department of Molecular and Cellular Physiology, Research Institute for Cell Design Medical Science, Graduate School of Medicine, Yamaguchi University, Ube City, Japan.
4. Department of Otolaryngology, Kobe City Medical Center General Hospital, Kobe city, Japan.