1. Usher Institute of Population Health Sciences & Informatics, University of Edinburgh, Edinburgh, United Kingdom.
2. Centre for Immunity, Infection and Evolution, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom.
3. Nuffield Department of Clinical Medicine, University of Oxford, John Radcliffe Hospital, Oxford, United Kingdom.
4. Institute of Infection and Global Health, University of Liverpool, Liverpool, United Kingdom.
5. International Livestock Research Institute, Nairobi, Kenya.