1. Department of Microbiology, College of Veterinary Medicine and Adult Stem Cell Research Center, Seoul National University, Seoul, Republic of Korea.
2. These authors contributed equally to this work.
3. Laboratory of Stem Cell and Tumor Biology, Department of Veterinary Public Health, College of Veterinary Medicine and Adult Stem Cell Research Center, Seoul National University, Seoul, Republic of Korea.
4. KRF Zoonotic Disease Priority Research Institute, College of Veterinary Medicine and Adult Stem Cell Research Center, Seoul National University, Seoul, Republic of Korea.
5. Department of Veterinary Internal Medicine, College of Veterinary Medicine and Adult Stem Cell Research Center, Seoul National University, Seoul, Republic of Korea.