1. Department of Medical Genetics, University of Wisconsin-Madison, Madison, Wisconsin, USA.
2. McPherson Eye Research Institute, University of Wisconsin-Madison, Madison, Wisconsin, USA.
3. Department of Ophthalmology and Visual Sciences, University of Wisconsin-Madison, Madison, Wisconsin, USA.
4. Department of Ophthalmology and Duke University Medical Center, Durham, North Carolina, USA.
5. Department of Cell Biology, Duke University Medical Center, Durham, North Carolina, USA.