1. Bioresource Center, RIKEN, Tsukuba, Ibaraki, Japan; CREST, JST, Kawaguchi, Saitama, Japan
2. Bioresource Center, RIKEN, Tsukuba, Ibaraki, Japan
3. CREST, JST, Kawaguchi, Saitama, Japan; Gene Research Center, Tokyo Institute of Technology, Yokohama, Kanagawa, Japan