1. Department of Antibody Drug Development, Tohoku University Graduate School of Medicine, Sendai, Japan.
2. New Industry Creation Hatchery Center, Tohoku University, Sendai, Japan.
3. Laboratory of Veterinary Pathology, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan.
4. Laboratory of Veterinary Surgery, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan.