1. Wellcome Centre Human Genetics, University of Oxford, Oxford, United Kingdom.
2. Laboratoire MIVEGEC (UMR CNRS 5290, UR IRD 224, UM), Montpellier, France.
3. National Institute for Health Research (NIHR) Nottingham Biomedical Research Centre, Nottingham University Hospitals NHS Trust and University of Nottingham, Nottingham, United Kingdom.
4. Nuffield Department of Medicine and the Oxford NIHR BRC, University of Oxford, Oxford, United Kingdom.
5. Centre for Virus Research, MRC-University of Glasgow, Glasgow, United Kingdom.