1. Department of Genetics and Genome Biology, University of Leicester, University Road, Leicester, United Kingdom.
2. Warwick Medical School, University of Warwick, Coventry, United Kingdom.
3. School of Life Sciences, University of Warwick, Coventry, United Kingdom.
4. School of Biosciences, University of Nottingham, Sutton Bonington Campus, Sutton Bonington, United Kingdom.