1. Centre for Early Human Development, Monash Institute of Reproduction and Development, Monash University, Clayton, Victoria, Australia
2. Centre for Early Human Development, Monash Institute of Reproduction and Development, Monash University, Clayton, Victoria, Australia and Genetics Australia Co-operative Ltd., Bacchus Marsh, Victoria, Australia
3. Department of Anatomy and Physiology, Royal Veterinary and Agricultural University, Frederiksberg, Denmark