1. Division of Biological Science, Graduate School of Science, Nagoya University, Chikusa-ku, Nagoya, Japan
2. Division of Applied Biosciences, Graduate School of Agriculture, Kyoto University, Sakyo-ku, Kyoto, Japan
3. Division of Biological Science, Graduate School of Science and Laboratory of Freshwater Fish Stocks, Bioscience Center, Nagoya University, Chikusa-ku, Nagoya, Japan