1. Department of Pathology, National Institute of Infectious Diseases, Tokyo, Japan; CREST, JST, Japan; Department of Medical Microbiology, Conway Institute of Biomolecular and Biomedical Research, University College Dublin, Belfield, Ireland
2. Department of Veterinary Science, National Institute of Infectious Diseases, Tokyo, Japan
3. Department of Pathology, National Institute of Infectious Diseases, Tokyo, Japan
4. Department of Pathology, National Institute of Infectious Diseases, Tokyo, Japan; CREST, JST, Japan
5. Department of Molecular Medicine, Institute for Molecular and Cellular Regulation, Gunma University, Gunma, Japan. 6Laboratory of Molecular and Cellular Pathology, Hokkaido University School of Medicine, Hokkaido, Japan
6. Department of Medical Microbiology, Conway Institute of Biomolecular and Biomedical Research, University College Dublin, Belfield, Ireland
7. CREST, JST, Japan; Laboratory of Molecular and Cellular Pathology, Hokkaido University of Medicine, Hokkaido, Japan