1. Division of Marine Life Science, Laboratory of Aquaculture Genetics and Genomics, Faculty and Graduate School of Fisheries Sciences, Hokkaido University, Hakodate, Japan.
2. Research Fellow of Japan Society for the Promotion of Science, Hokkaido University, Nanae, Japan.
3. Nanae Fresh-Water Station, Field Science Center for Northern Biosphere, Hokkaido University, Nanae, Japan.