1. Research Center for Plant Conservation, Botanic Gardens and Forestry, National Research and Innovation Agency, Cibinong 16911, Indonesia
2. Faculty of Biology, Gadjah Mada University, Sekip Utara Yogyakarta 55281, Indonesia
3. Research Center for Horticulture and Plantation, National Research and Innovation Agency, Cibinong 16911, Indonesia
4. Departement of Conservation, PT Mitra Natura Raya, Bogor Botanical Garden, Bogor 16122, Indonesia