1. Department of Plant Science, College of Agriculture and Life Sciences, Seoul National University, Seoul 08826, Korea
2. Institutes of Green-Bio Science and Technology, Seoul National University, Pyeongchang 25354, Korea
3. Graduate School of International Agricultural Technology, Seoul National University, Pyeongchang 25354, Korea
4. Radiation Breeding Research Team, Advanced Radiation Technology Institute, Korea Atomic Energy Research Institute, Jeongeup 56212, Korea