1. Department of Life Sciences (Insects), The Natural History Museum, Cromwell Road, London, SW7 5BD, United Kingdom;
2. USDA/ARS Systematic Entomology Laboratory, c/o National Museum of Natural History, MRC-168, 10th St. and Constitution Ave., Smithsonian Institution, Washington, DC 20560, USA;
3. Department of Entomology, National Museum, Cirkusová 1740, CZ-193 00 Praha 9—Horní Počernice, Czech Republic;