1. Department of Botany, University of Wisconsin – Madison, WI, 53706 (United States)
2. Department of Ecology, Evolution, and Organismal Biology, Iowa State University, Ames, IA, 50011 (United States)
3. Department of Ecology, Evolution, and Organismal Biology, Iowa State University, Ames, IA, 50011 (United States) and Department of Biology, University of Massachusetts, 611 North Pleasant Street, Amherst, MA 01003 (United States)
4. Department of Botany, University of Wisconsin – Madison, WI, 53706 (United States) and Wisconsin Institute for Discovery, 330 N Orchard St, Madison, WI 53715 (United States)