1. Department of fundamental sciences, University of Quebec in Chicoutimi, Quebec (Canada)
2. AMAP, Univ. Montpellier, CIRAD, CNRS, INRA, IRD, F-34098 Montpellier (France) and Department of Ecology and Evolutionary Biology, and Natural History Museum and Biodiversity Institute, University of Kansas, Lawrence, KS 66045-7534 (United States)
3. AMAP, Univ. Montpellier, CIRAD, CNRS, INRA, IRD, F-34098 Montpellier (France)