1. CR2P (CNRS, MNHN, Sorbonne Université), Muséum national d'Histoire naturelle, case postale 48, 57 rue Cuvier, F-75231 Paris cedex 05 (France)
2. Evolutionary Studies Institute, Witwatersrand University, 1 Jan Smuts Ave., Braamfontein, Johannesburg 2000, South Africa