1. UMR ISyEB-7205 (CNRS, MNHN, UPMC, EPHE), Museum National d'Histoire Naturelle, 57 Rue Cuvier, CP 39, 75231 cedex Paris 05 (France)
2. Department of Biology, Norwegian University of Science and Technology, 7491 Trondheim (Norway)
3. Department of Botany, The Field Museum, 1400 South Lake Shore Drive, Chicago, IL 60605–2496 (United States)
4. UMS PatriNat-2006 (OFB, CNRS, MNHN), Museum National d'Histoire Naturelle, 57 Rue Cuvier, CP 39, 75231 cedex Paris 05 (France)