1. Department of Biology, University of Alabama at Birmingham, Birmingham, AL 35294 (United States)
2. GEOMAR Helmholtz-Zentrum für Ozeanforschung Kiel, Düsternbrooker Weg 20, D-23105 Kiel (Germany)
3. College of Marine Life Sciences, Ocean University of China, 5 Yushan Road, Qingdao 266003 (China) and Institute of Evolution and Marine Biodiversity, Ocean University of China, Qingdao 266003 (China)