1. G.T. Velasquez Phycological Herbarium and The Marine Science Institute, College of Science, University of the Philippines Diliman, Quezon City, 1101 (Philippines)
2. Department of Natural History Sciences, Graduate School of Science, Hokkaido University, Sapporo, 060-0810 (Japan) and Research Center for Inland Seas, Kobe University, Rokkodai 1-1, Nadaku, Kobe 657-8501 (Japan)
3. Department of Biological Sciences, Faculty of Science, Hokkaido University, Sapporo, 060-0810 (Japan)