1. Central Department of Zoology, Institute of Science and Technology, Tribhuvan University, Kathmandu 44618, Nepal
2. Washington National Primate Research Center, University of Washington, Seattle, WA 98195, USA.
3. Departments of Psychology, Global Health, and Anthropology, Center for Global Field Study, and Washington National Primate Research Center, University of Washington, Seattle, WA 98195, USA.