1. Pablo de Olavide University, ctra. de Utrera, km 1, Ed. 22 41013, Seville, Spain
2. Department of Integrative Ecology, Estación Biológica de Doñana (EBD‐CSIC), Avda. Americo Vespucio s/n 41092, Seville, Spain
3. Departamento de Biología Vegetal y Ecología, Universidad de Sevilla, Apartado 1095 41080, Sevilla, Spain
4. McGill University, Plant Science, Macdonald Campus 21111 Lakeshore, Ste. Anne de Bellevue, Quebec H9X 3V9, Canada
5. The Morton Arboretum, 4100 Illinois Route 53, Lisle, Illinois 60532 USA
6. Botany Department, Field Museum of Natural History, Chicago, Illinois 60605 USA