1. Florida Museum of Natural History, University of Florida, Gainesville, Florida 32611, USA
2. Department of Biology, University of Florida, Gainesville, Florida 32611, USA
3. Royal Botanic Gardens, Kew, Natural Capital and Plant Health Department, Richmond, Surrey TW9 3DS, UK
4. Kangwon National University, Department of Applied Plant Sciences, Chuncheon 24341, Korea
5. Genetics Institute, University of Florida, Gainesville, Florida 32610, USA