1. Department of Evolution, Ecology, and Organismal Biology, The Ohio State University, 318 West 12th Avenue, Columbus, Ohio 43210 USA;
2. Department of Ecology and Evolutionary Biology, University of Kansas, 1200 Sunnyside Ave, Lawrence, Kansas 66045 USA; and
3. Department of Biological Sciences, California State University Sacramento, 6000 J Street, Sacramento, California 95819 USA