1. 2Department of Botany and Plants Sciences, University of California, Riverside, California 92521 USA;
2. 3Department of Ecology and Evolutionary Biology, University of California, Irvine, California 92697 USA;
3. 4Jardín de Aclimatación de La Orotava, Calle Retama Num. 2, 38400, Puerto de La Cruz, Tenerife, Canary Islands, Spain