1. 2Laboratory of Anatomy and Morphology, V. L. Komarov Botanical Institute of Russian Academy of Sciences, Prof. Popov Street 2, 197376, St. Petersburg, Russia;
2. 3School of Biological Sciences, Washington State University, Pullman, Washington 99164‐4236 USA;
3. 4Department of Biology, Faculty of Science, University of Tehran, P.O. Box 14155‐6455, Tehran, Iran