1. La Selva Biological Station, Organization for Tropical Studies, Interlink 341, P.O. Box 02‐5635 Miami, Florida 33102‐5635 USA;
2. Organismic and Evolutionary Biology, Harvard University, 3119 Biological Laboratories, 16 Divinity Avenue, Cambridge, Massachusetts 02138 USA;
3. Institute of Pacific Islands Forestry, 1151 Punchbowl Street, Suite 323, Honolulu, Hawaii 96813 USA