1. The Ohio State University Herbarium, Department of Evolution, Ecology, and Organismal Biology, 1315 Kinnear Road, Columbus, Ohio 43212 USA
2. Institute of Arctic Biology, 205 West Research Building, University of Alaska‐Fairbanks, Fairbanks, Alaska 99775 USA
3. Institute of Ecology and Earth Sciences, University of Tartu, 40 Lai Street, Tartu 51005 Estonia