1. Department of Natural Resource Management, South Dakota State University, 1390 College Avenue, Brookings, South Dakota 57007 USA
2. Department of Biological Sciences, University of Idaho, 875 Perimeter Drive, MS 3051, Moscow, Idaho 83844‐3051 USA
3. Stillinger Herbarium, University of Idaho, 875 Perimeter Drive, MS 1133, Moscow, Idaho 83844‐1133 USA
4. Institute for Bioinformatics and Evolutionary Studies (IBEST), University of Idaho, 875 Perimeter Drive, MS 3051, Moscow, Idaho 83844‐3051 USA
5. Department of Ecology and Evolutionary Biology, University of Michigan, 830 North University Avenue, Ann Arbor, Michigan 48109 USA
6. Australian National Herbarium, CSIRO National Research Collections, Canberra, Australia