1. Department of Botany, The Field Museum, 1400 South Lake Shore Drive, Chicago, Illinois 60605 USA
2. Naturhistoriska Riksmuseet, Enheten för Kryptogambotanik, Box 50007 104 05 Stockholm, Sweden
3. Fundación Charles Darwin, Casilla 200144, Isla Santa Cruz, Galapagos, Ecuador
4. Department of Biology, Duke University, 125 Science Drive, Durham, North Carolina 27708, USA