1. Department of Ecology and Evolution, University of Lausanne, 1015 Lausanne, Switzerland
2. Department of Computational Biology, University of Lausanne, 1015 Lausanne, Switzerland
3. Swiss Institute of Bioinformatics, Quartier Sorge, 1015 Lausanne, Switzerland
4. Conservatoire et Jardin botaniques de la Ville de Genève and Laboratory of Plant Systematics and Biodiversity, University of Geneva, Chemin de l'Impératrice 1, 1292 Chambésy, Geneva, Switzerland